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List of 12-letter words containing •••••

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There are 20 twelve-letter words containing E, R, S and 3U

ACUPUNCTURES• acupunctures n. Plural of acupuncture.
• acupunctures v. Third-person singular simple present indicative form of acupuncture.
• ACUPUNCTURE n. the complementary therapy of puncturing the skin with needles at specified points in order to cure illness.
AQUACULTURES• aquacultures n. Plural of aquaculture.
• AQUACULTURE n. the practice of using the sea, lakes etc. for fish-farming, also AQUICULTURE.
AQUICULTURES• aquicultures n. Plural of aquiculture.
• AQUICULTURE n. the practice of using the sea, lakes etc. for fish-farming, also AQUACULTURE.
CREPUSCULOUS• crepusculous adj. Alternative form of crepuscular.
• CREPUSCULOUS adj. relating to twilight, also CREPUSCULAR.
CUPULIFEROUS• cupuliferous adj. (Botany) Bearing, or furnished with, a cupule.
• CUPULIFEROUS adj. bearing cupules.
EQUINUMEROUS• equinumerous adj. (Mathematics) Having equal cardinality.
• EQUINUMEROUS adj. having the same number of members.
FURFURACEOUS• furfuraceous adj. Made of bran.
• furfuraceous adj. Resembling bran; having or characterised by small flakes that resemble bran; resembling dandruff.
• FURFURACEOUS adj. containing bran or particles resembling bran, also FURFUROUS.
FURUNCULOSES• furunculoses n. Plural of furunculosis.
• FURUNCULOSIS n. an affliction of boils.
GROUPUSCULES• groupuscules n. Plural of groupuscule.
• GROUPUSCULE n. a small clique or faction.
MUSCULATURES• musculatures n. Plural of musculature.
• MUSCULATURE n. the muscles of all or a part of the animal body.
NEUROHUMOURS• neurohumours n. Plural of neurohumour.
• NEUROHUMOUR n. a neurohormone, esp. a neurotransmitter, also NEUROHUMOR.
PRESUMPTUOUS• presumptuous adj. Going beyond what is right, proper, or appropriate because of an excess of self-confidence or arrogance.
• PRESUMPTUOUS adj. overstepping due bounds (as of propriety or courtesy).
RANUNCULUSES• ranunculuses n. Plural of ranunculus.
• RANUNCULUS n. a genus of herbs, mostly with yellow flowers, including crowfoot and buttercups.
SUBOPERCULUM• suboperculum n. (Anatomy) The lower opercular bone in fishes.
• SUBOPERCULUM n. in fishes, a bone below and partly behind the gill cover.
SUBSTRUCTURE• substructure n. The supporting part of a structure (either physical or organizational; the foundation).
• substructure n. The earth or gravel that the railway tracks are embedded in.
• substructure n. (Marxism) Synonym of base (“the forces and relations of production that produce the necessities and…
SUPERNACULUM• supernaculum adv. (Obsolete) According to the rules of an old drinking game in which the drinker upturned the empty cup…
• supernaculum adv. To the last drop, to the bottom.
• supernaculum n. (Obsolete) Excellent wine that one would wish to drink to the last drop.
UNCOURAGEOUS• uncourageous adj. Lacking courage.
• UNCOURAGEOUS adj. not courageous.
UNSCULPTURED• unsculptured adj. Not sculptured.
• UNSCULPTURED adj. not sculptured.
UNSTRUCTURED• unstructured adj. Lacking structure.
• UNSTRUCTURED adj. lacking structure or organization.
USURIOUSNESS• usuriousness n. Quality of being usurious.
• USURIOUSNESS n. the state of being usurious.

Scrabble words — in black are valid world wide — in RED are not valid in North America — in GREEN are valid only in North America.
Definitions are short excerpt from the WikWik.org and 1Word.ws.

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